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Scientific Publications by Stephen Mifsud
   

This page lists the publications written by Stephen Mifsud (author of MaltaWildPlants.com) related with his botanical research on the Maltese Islands. Many of his publications are published in peer-reviewed scientific journals but certain research is suitable to be published online on this dedicated page, hence serving as a communication to the scientific community, students and general public.

These online scientific communications are digital publications which allows large number of images, tables, video and other forms of digital data. It helps readers to determine and gauge the validity of these online articles much better. The advantages of online publishing are: fast to publish, reaching a much broader spectrum of e-readers, indexed in search engines like Google and Yahoo, easy to access through internet, free of charge, easy to update and finally, users can easily send their feedback and criticism to the author.

This page is divided into two main categories: 1) printed (bibliographical) papers published in scientific journals and 2) the online articles that are supplied through links in this page. These can also be referenced in one's work and papers. The former are not available online due to copyright restrictions, but can purchased from the corresponding journal. Hope that this page will meet its goal of communicating important new scientific data which can be of use by the scientist, student and general public.

Page last modified on:   24 Apr 2015 (v: 2.3)


TABLE OF ARTICLE TITLES
Online Articles of Scientific Value
Link Title / Authors / Date Link Title / Authors / Date
W01 A FLORISTIC SURVEY ON THE GOZITAN ISLETS OF TAC-CAWL AND TAL-HALFA IN THE MALTESE ISLANDS
Stephen Mifsud (2011)
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Printed Scientific Papers
Link Title / Authors / Date Link Title / Authors / Date
P01 Two hybrid orchids described from Malta: Ophrys x tumentia nothosubsp. tumentia and Ophrys x tumentia nothosubsp. sancti-martinii
Stephen Mifsud (Dec 2014)
P02 Promotion of the Maltese Ophrys bombyliflora forma parviflora to a variety
Stephen Mifsud (Dec 2014)
P03 Chromosome number of Elatine gussonei (Sommier) Brullo (Elatinaceae) and its distribution on the Maltese Islands
Anna Kalinka, Stephen Mifsud, Agnieska Popiela, Magdalena Achremi (2014)
P04 A study of the genus Persicaria Miller (Polygonaceae) in the Maltese Islands
Stephen Mifsud (2014)
P05 Bromus cathatrticus Vahl (Fam. Poaceae), A new record for the Maltese Islands
Timothy J. Tabone, Alex Casha & Stephen Mifsud (2014)
P06 Marasmius corbariensis (Roum.) Singer (Family Marasmiaceae Roze Ex Kuhner 1980) – A new fungal species for Malta
Stephen Mifsud (2014)
P07 Recent records of spiders (Arachnida: Araneae) from Malta
David Dandria, Alfred Gatt Floridia & Stephen Mifsud (2014)
P08 Polypodium vulgare L. subsp. melitense new subspecies from Gozo, Maltese Islands (Pteridophyta: Polypodiaceae Bercht. & J. Presl.)
Adalberto Peroni, Gabriele Peroni & Stephen Mifsud (2013)
P9 Preliminary observations from long-term studies of Gladiolus l. (Iridaceae) for the Maltese Islands
Stephen Mifsud and Anthony P. Hamilton (Aug 2013)
P10 Althenia orientalis (Tzvelev) Garcia-Mur. & Talavera (Zanichelliaceae Dum.): An addition to the Maltese Flora (Central Mediterranean)
Alex Casha & Stephen Mifsud (Aug 2013)
P11 Distribution of some rare or endemic chasmophytic and rupestral species growing along the coastal cliffs of the Maltese Islands
Stephen Mifsud (Aug 2013)
P12 Recent observations on the Ophrys of Malta and Gozo: addendum
Stephen Mifsud & Leslie Lewis (May 2013)
P13 Fumana laevis (Cav.) Pau (Cistaceae): New to the flora of Malta and considerations on Fumana (Dunal) Spach from the Maltese Flora
Stephen Mifsud (May 2012)
P14 Recent observations on Ophrys species and hybrids of the Maltese archipelago
Stephen Mifsud & Leslie Lewis (Nov 2011)
P15 New records of the genus Pachygrapsus (Crustacea: Decapoda) from the Central Mediterranean sea with a review of its Mediterranean zoogeography
Fabio Crocetta, S. Mifsud, P. Paolini, J. Piscopo & P. J. Schembri (2011)
P16 A new record of Serapias bergonii [Fam. Orchidaceae] from the Maltese Islands
Stephen Mifsud (2010)
P17 Two new alien grasses from sand gunes Of Ghadira bay in Malta
Stephen Mifsud & Alex Casha (2010)
P18 First occurences of Lemna minuta [Fam. Lemnaceae] in the Maltese islands
Stephen Mifsud (2010)
P19 Records of Narcissus elegans [Fam. Amaryillidaceae] and notes on the wild Narcissus in the Maltese islands
Stephen Mifsud & Emanuel Caruana (2010)
P20 A new form of Ophrys bombyliflora link described from Malta
Stephen Mifsud (2009)
P21 Rediscovery and present distribution of Valantia hispida L. (Fam. Rubiaceae) in the Maltese Islands
Stephen Mifsud (2009)
P22 Update of the Maltese Flora (Central Mediterranean) including very rare species or species thought to be extinct from mainland Malta or its islands
Stephen Mifsud (2009)
P23 Zannichellia major Boenn. ex Rchb. - a first record of a non-endemic species of Zannichellia l. (Fam. Zannichelliaceae Dum.) from the Maltese Islands
Stephen Mifsud (2009)
P24 Four new records for the flora of the Maltese Islands - Dactyloctenium aegyptium (L.) P.Beauv. (Fam. Poaceae), Amaranthus muricatus (Gillies Ex Moq.) Hieron. (Fam. Amaranthaceae), Fumaria reuteri Boiss.(Fam. Fumariaceae) and Spirodela oligorrhiza(Kurtz) Hegelm. (Fam. Lemnaceae)
Stephen Mifsud (2008)
P25 An assessment on the distribution of Iris pseudopumila Tineo [Fam. Iridaceae] in the Maltese Islands and comparison of its two flower forms
Stephen Mifsud (2008)
P26 Three new species from the genus Ophrys, section Pseudophrys [Fam: Orchidaceae] on the Maltese Islands
Stephen Mifsud (2008)
P27 Updates in the Flora of the Maltese Islands (Central Mediterranean)
Stephen Mifsud (2007)
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Printed Scientific Papers


Paper 01 Updates in the Flora of the Maltese Islands (Central Mediterranean)
Author(s) / Yr. of Publ. Stephen Mifsud (2007)
Abstract Discovery of new populations in Malta of the very rare Ornithogalum divergens, Asparagus stipularis, Muscari commutatum, Muscari neglectum, Vicia bithynica, Cardaria draba and Abutilion theophrasti, rediscovery of Trifolium pratense (last report of this species 80 years ago) and a new species from Malta - Calendula bicolor
Keywords Muscari commutatum / Muscari neglectum / Calendula bicolor / Linaria pseudolaxiflora / Asparagus stipularis / Ornithogalum divergens / Vicia bithynica / Cardaria draba / Abutilion theophrasti / Trifolium pratense / rediscovery / rare species / Malta / Gozo / flora
Citation Mifsud, S. (2007): Updates in the Flora of the Maltese Islands (Central Mediterranean).   The Central Mediterranean Naturalist, 4(3): 171-179. Nature Trust, Malta.
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Paper 02 Three new species from the genus Ophrys, section Pseudophrys [Fam: Orchidaceae] on the Maltese Islands
Author(s) / Yr. of Publ. Stephen Mifsud (2008)
Abstract This article deals with the first substantiated records from the Maltese islands of Ophrys lupercalis J. Devillers-Terschuren & P. Devillers and Ophrys lojaconoi P. Delforge, both forming part of the Ophrys fusca group, and of Ophrys phryganae J. Devillers-Terschuren & P. Devillers, belonging to the Ophrys lutea group. Identification is based on isometric morphological analysis, following the Delforge taxonomical system. Treatment of these taxa according to another taxonomical system by Baumann, Kunkele and Lorenz is also discussed.
Keywords Ophrys lojaconoi / Ophrys lupercalis / Ophrys phryganae / Ophrys fusca group / Ophrys lutea group / Ophrys / Orchidaceaea / Malta
Citation Mifsud, S. (2008): Three new species from the genus Ophrys, section Pseudophrys [Fam: Orchidaceae] on the Maltese Islands.   The Central Mediterranean Naturalist, 4(4): 243-252. Nature Trust, Malta
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Paper 03 An assessment on the distribution of Iris pseudopumila Tineo [Fam. Iridaceae] in the Maltese Islands and comparison of its two flower forms
Author(s) / Yr. of Publ. Stephen Mifsud (2008)
Abstract The distribution of Iris pseudopumila Tineo in the Maltese islands is given, including information on four new populations. A comparison study between three populations of the yellow-flower form, and between the violet and yellow forms are provided, the latter showing marked morphological differences between the two colour forms in the Maltese islands.
Keywords Iris pseudopumila / Iridaceae / distribution / colour forms / Malta.
Citation Mifsud, S. (2008): An assessment on the distribution of Iris pseudopumila Tineo [Fam. Iridaceae] in the Maltese Islands and comparison of its two flower forms.   The Central Mediterranean Naturalist, 4(4): 253-262. Nature Trust, Malta
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Paper 04 Four new records for the flora of the Maltese Islands - Dactyloctenium aegyptium (L.) P.Beauv. (Fam. Poaceae), Amaranthus muricatus (Gillies Ex Moq.) Hieron. (Fam. Amaranthaceae), Fumaria reuteri Boiss.(Fam. Fumariaceae) and Spirodela oligorrhiza(Kurtz) Hegelm. (Fam. Lemnaceae)
Author(s) / Yr. of Publ. Stephen Mifsud (2008)
Abstract Four plant species are reported as new to the flora of the Maltese islands: Dactyloctenium aegyptium (L.) P.Beauv. (Family Poaceae), Amaranthus muricatus (Gillies ex Moq.) Hieron. (Family Amaranthaceae), Fumaria reuteri Boiss. (Family Fumariaceae) and Spirodela oligorrhiza (Kurtz) Hegelm. (Family Lemnaceae). The article gives worldwide distribution, plant characteristics and information on the location and habitat that they were found in. The article is complemented by several illustrations
Keywords Dactyloctenium aegyptium / Amaranthus muricatus / Fumaria reuteri / Spirodela oligorrhiza / new records / Malta / flora
Citation Mifsud, S. (2008): Four new records for the flora of the Maltese Islands - Dactyloctenium aegyptium (L.) P.Beauv. (Fam. Poaceae), Amaranthus muricatus (Gillies Ex Moq.) Hieron. (Fam. Amaranthaceae), Fumaria reuteri Boiss.(Fam. Fumariaceae) and Spirodela oligorrhiza(Kurtz) Hegelm. (Fam. Lemnaceae).   The Central Mediterranean Naturalist, 4(4): 263-273. Nature Trust, Malta
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Paper 05 Zannichellia major Boenn. ex Rchb. - a first record of a non-endemic species of Zannichellia l. (Fam. Zannichelliaceae Dum.) from the Maltese Islands
Author(s) / Yr. of Publ. Stephen Mifsud (2009)
Abstract The first record of Zannichellia major Boenn. ex Rchb. is reported from the Maltese islands. Detailed morphological comparison of this population with the endemic and frequent Zannichellia melitensis Brullo, Giusso & Lanfranco and other monoecious Zannichellia species is given. Distribution of this species is discussed.
Keywords Zannichellia major / Zannichellia palustris / Zannichellia melitensis / Maltese islands, Wetlands / Aquatic flora / Mediterranean region / Malta
Citation Mifsud, S. (2009): Zannichellia major Boenn. ex Rchb. - a first record of a non-endemic species of Zannichellia l. (Fam. Zannichelliaceae Dum.) from the Maltese Islands.   The Central Mediterranean Naturalist, 5(1): 1-6 Nature Trust, Malta
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Paper 06 Update of the Maltese Flora (Central Mediterranean) including very rare species or species thought to be extinct from mainland Malta or its islands
Author(s) / Yr. of Publ. Stephen Mifsud (2009)
Abstract Rediscovery of Lotus halophilus from mainland Malta (presumed to be found only in the island of Comino), Xanthium spinosum (presumed extinct), first record of Myrtus commuis from the island of Gozo and the very rare plant Asplenium trcihomanes are reported from the Maltese Islands. An account about the each population and an updated distribution of the species in Malta is given.
Keywords Lotus halophilus / Myrtus communis / Xanthium spinosum / Asplenium trichomanes / rare species / extinct / rediscovery / Malta / flora
Citation Mifsud, S. (2009): Update of the Maltese Flora (Central Mediterranean) including very rare species or species thought to be extinct from mainland Malta or its islands.   The Central Mediterranean Naturalist, 5(1): 7-16 Nature Trust, Malta
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Paper 07 Rediscovery and present distribution of Valantia hispida L. (Fam. Rubiaceae) in the Maltese Islands
Author(s) / Yr. of Publ. Stephen Mifsud (2009)
Abstract Valantia hispida L. is an indigenous species presumed extinct from the Maltese islands due to lack of any substantiated records for more than 130 years. The rediscovery of this species is reported. An account of the population found in the South East area of the island of Gozo is given. Morphological differences between V. hispida L. and the frequent V. muralis L. are illustrated. The taxon V. muralis var. hirsuta reported from Malta in old literature is discussed.
Keywords Valantia hispida / extinct / rediscovery / Mediterranean, Malta / Gozo / flora
Citation Mifsud, S. (2009): Rediscovery and present distribution of Valantia hispida L. (Fam. Rubiaceae) in the Maltese Islands.   The Central Mediterranean Naturalist, 5(1): 17-23. Nature Trust, Malta
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Paper 08 A new form of Ophrys bombyliflora link described from Malta
Author(s) / Yr. of Publ. Stephen Mifsud (2009)
Abstract A new form, Ophrys bombyliflora Link forma parviflora S. Mifsud is described for plants with significantly smaller lips from the typical form of O. bombyliflora and with a bright, contrasting speculum. These plants were observed as homogenous populations in 2 different sites in Malta, isolated from populations of the locally common Ophrys bombyliflora Link.
Keywords Orchidaceae / Ophrys bombyliflora forma parviflora / Ophrys / taxonomy / Malta / new taxon
Citation Mifsud, S. (2009): A new form of Ophrys bombyliflora link described from Malta.   Journal of European Orchids, 41(3/4): 611-622, December 2009, Germany
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Paper 9 Records of Narcissus elegans [Fam. Amaryillidaceae] and notes on the wild Narcissus in the Maltese islands
Author(s) / Yr. of Publ. Stephen Mifsud & Emanuel Caruana (2010)
Abstract Substantiated records of Narcissus elegans are reported from Malta and for the first time in Gozo. Habitat and ecological analysis of the three different populations and a distribution map is given. Morphological characters between the populations are compared between each other and the described taxa. Also discussed and illustrated are the intermediate range of specimens found in Malta between N. serotinus, N. elegans and N. tazetta
Keywords Narcissus elegans / Narcissus serotinus / Narcissus tazetta / new records / Narcissus intermediates / Malta / Gozo
Citation Mifsud, S. & Caruana, E. (2010): Records of Narcissus elegans [Fam. Amaryillidaceae] and notes on the wild Narcissus in the Maltese islands.   The Central Mediterranean Naturalist, 5(2): 19-29. Nature Trust, Malta
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Paper 10 First occurences of Lemna minuta [Fam. Lemnaceae] in the Maltese islands
Author(s) / Yr. of Publ. Stephen Mifsud (2010)
Abstract The existence of Lemna minuta Kunth is confirmed from two sites in mainland Malta and one in the island of Gozo and reported for the first time in the Maltese islands. Notes on its historical and current distribution in Europe, its invasive properties and its habitat preferences in Malta are given. Also included are the distinguishing morphological features from the closely related Lemna minor L., another frequent alien species of ponds and valleys in the Maltese islands.
Keywords Lemna minuta / valleys / watercourses / alien species / invasive species / flora of Malta
Citation Mifsud, S. (2010): First occurences of Lemna minuta [Fam. Lemnaceae] in the Maltese islands.   The Central Mediterranean Naturalist, 5(2): 1-4. Nature Trust, Malta
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Paper 11 Two new alien grasses from sand gunes Of Ghadira bay in Malta
Author(s) / Yr. of Publ. Stephen Mifsud & Alex Casha (2010)
Abstract Cenchrus echinatus L., a new species for the Maltese Islands, and Dactyloctenium aegyptium, only recorded from the island of Gozo, has been found at the remnant sand dunes of l-G?adira bay, a popular sandy bay located in Mellieha situated in the North of mainland Malta. Their invasive reputation and their presumed introduction routes to our islands are discussed.
Keywords Cenchrus echinatus / Dactyloctenium aegyptium / sand dunes / coasts / alien species / invasive species / flora of Malta
Citation Mifsud, S. & Casha, A. (2010): Two new alien grasses from sand gunes Of Ghadira bay in Malta.   The Central Mediterranean Naturalist, 5(2): 6-9. Nature Trust, Malta
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Paper 12 A new record of Serapias bergonii [Fam. Orchidaceae] from the Maltese Islands
Author(s) / Yr. of Publ. Stephen Mifsud (2010)
Abstract Serapias bergonii E.G.Camus is a very rare species for the Maltese islands and was assumed extinct from the island of Gozo due to lack of substantiated records. This paper gives details about a new record from Xlendi, a locality on the West coast of the island of Gozo.
Keywords Serapias bergonii / Serapias vomeracea subsp. laxiflora / Orchidaceae / Mediterranean region / Malta / Gozo
Citation Mifsud, S. (2010): A new record of Serapias bergonii [Fam. Orchidaceae] from the Maltese Islands.   The Central Mediterranean Naturalist, 5(2): 11-17. Nature Trust, Malta
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Paper 13 New records of the genus Pachygrapsus (Crustacea: Decapoda) from the Central Mediterranean sea with a review of its Mediterranean zoogeography
Author(s) / Yr. of Publ. Fabio Crocetta, S. Mifsud, P. Paolini, J. Piscopo & P. J. Schembri (2011)
Abstract The occurrence of Pachygrapsus maurus and Pachygrapsus transversus is reported from the Maltese Islands for the first time on the basis of one specimen of P. maurus collected in 1990 and numerous recent specimens, and the distribution of the two species is mapped. The controversial presence of P. maurus in Italy is confirmed and two new sites for this species are reported, including the first for the mainland of Italy. The examination of the historical specimen of P. maurus from the Genova area revealed a misidentification of P. transversus; this record could be a result of ship-mediated transport. First notes on the habitat of P. maurus in the central Mediterranean Sea are given. Updated maps of the distribution of P. maurus and P. transversus in the Mediterranean are provided and the zoogeography of these species is revisited.
Keywords Pachygrapsus marmoratus / Pachygrapsus maurus / Pachygrapsus transversus / Zoogeography / Mediterranean Sea.
Citation F. Crocetta, S. Mifsud, P. Paolini, J. Piscopo and P. J. Schembri (2011): New records of the genus Pachygrapsus (Crustacea: Decapoda) from the Central Mediterranean sea with a review of its Mediterranean zoogeography.   Mediterranean Marine Science, Medit. Mar. Sci., 12/1, 2011
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Paper 14 Recent observations on Ophrys species and hybrids of the Maltese archipelago
Author(s) / Yr. of Publ. Stephen Mifsud & Leslie Lewis (Nov 2011)
Abstract Mifsud, S. & L. Lewis (2011): Recent Observations on Ophrys species and hybrids of the Maltese Archipelago.- J. Eur. Orch. 43 (3): 609-650. Detailed field studies carried out between late December 2007 and early April 2011 show that the Ophrys iricolor s.l. group on Malta comprises two morphologically distinct forms which flower at different times in two main waves, although some intermediates also occur. The first form flowers from early December to late January and has a medium sized labellum with an underlip which is entirely green or is stained pale red without a clear–cut yellowish–green border. The second form flowers from late February to early April and has a longer labellum with an underlip which is normally stained red with a clear–cut yellowish–green border. It is concluded that the first form is attributable to O. mesaritica, consistent with the existing records of this species on the Maltese archipelago, whereas the second form is attributable to O. vallesiana. It is also concluded that some intermediate plants, especially those flowering in mid–February between the two main flowering waves, are probably O. mesaritica introgressed/hybridised with O. vallesiana (that is O. mesaritica × O. vallesiana).


Field studies of Ophrys caesiella at various locations on Malta show that this species is more variable than was originally described by DELFORGE (2000) and support opinions that O. caesiella and O. gazella are the same species. In addition, an Ophrys similar to O. caesiella but possessing prominent basal ridges and a geniculated labellum, initially identified as O. bilunulata × O. mesaritica by DELFORGE (1993) and subsequently incorrectly re–identified as O. pectus (DEVILLERS & DEVILLERS–TERSCHUREN 2000a), has been refound at its original location as well as elsewhere. It is concluded that it is the result of hybridization and/or introgression of O. caesiella with/by O. vallesiana (that is O. caesiella × O. vallesiana). However, the possibility cannot be excluded that the O. iricolor s.l. parent of some plants is instead O. mesaritica, or that some plants are complex hybrids of O. caesiella, O. vallesiana and O. mesaritica. Details of four Ophrys taxa (namely O. lojaconoi, O. lupercalis, O. phryganae and O. bombyliflora forma parviflora) recorded on the Maltese archipelago since the publication of a comprehensive list of Maltese orchids by BARTOLO et al. (2001) are also given, as well as new locations for some of the rarer previously listed species.
Keywords Orchidaceae; Ophrys caesiella, Ophrys gazella, Ophrys mesaritica, Ophrys pectus, Ophrys vallesiana, Ophrys caesiella × Ophrys vallesiana, Ophrys mesaritica × Ophrys vallesiana. Malta, Gozo.
Citation Mifsud, S. & Lewis, L. (Nov 2011): Recent observations on Ophrys species and hybrids of the Maltese archipelago.   Journal Europäischer Orchideen, 43 (3): 609-650. Germany.
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Paper 15 Fumana laevis (Cav.) Pau (Cistaceae): New to the flora of Malta and considerations on Fumana (Dunal) Spach from the Maltese Flora
Author(s) / Yr. of Publ. Stephen Mifsud (May 2012)
Abstract Fumana laevis (Cav.) Pau (Cistaceae): nuova per la Flora di Malta, e considerazioni su Fumana (Dunal) Spach per la Flora di Malta — Fumana laevis (Cav.) Pau è riconosciuta per la Flora di Malta. Vengono fornite informazioni sulla tassonomia di F. laevis in relazione alle altre tre specie del genere presenti nella fl ora di Malta. Viene inoltre discussa la sua distribuzione locale e generale, la sua ecologia e condizione attuale.
Keywords Cistaceae, Fumana, Fumana laevis, Flora, Maltese Islands
Citation Mifsud, S. (May 2012): Fumana laevis (Cav.) Pau (Cistaceae): New to the flora of Malta and considerations on Fumana (Dunal) Spach from the Maltese Flora.   Webbia, 67(1): 101-107. 2012
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Paper 16 Recent observations on the Ophrys of Malta and Gozo: addendum
Author(s) / Yr. of Publ. Stephen Mifsud & Leslie Lewis (May 2013)
Abstract As previously reported by the present authors (MIFSUD & LEWIS 2011), field studies at various locations on Malta indicate that Ophrys caesiella and O. gazella, and O. africana are the same species. The name O. caesiella is now chosen for this combined species. LOWE (2011) similarly concluded that O. caesiella, O. gazella and O. africana were synonyms but also further concluded that O. subfusca (Rchb. f.) Hausskn. is an earlier synonym for these taxa. However, in the opinion of the present authors, the assumptions on which Lowe's further conclusion is based cannot be validated and (if legitimate) O. subfusca should therefore continue to be considered a member of the O. lutea complex. New sites for O. caesiella on Malta and Gozo are also reported. DELFORGE (2012) concluded that the early-flowering form of O. iricolor s.l. on Malta was morphologically different from O. mesaritica on Crete and described it under the new name O. hospitalis. This is now renamed Ophrys iricolor subsp. hospitalis. Reference is also made to the legal protection of wild orchids on Malta.
Keywords Orchidaceae, Ophrys africana, Ophrys caesiella, Ophrys gazella, Ophrys hospitalis, Ophrys iricolor subsp. hospitalis, Ophrys lutea var. subfusca, Ophrys subfusca; legal protection of wild orchids.
Citation Mifsud, S. & Lewis, L. (May 2013): Recent observations on the Ophrys of Malta and Gozo: addendum .   Journal Europäischer Orchideen, 45 (1): 105-120, Germany
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Paper 17 Distribution of some rare or endemic chasmophytic and rupestral species growing along the coastal cliffs of the Maltese Islands
Author(s) / Yr. of Publ. Stephen Mifsud (Aug 2013)
Abstract This paper gives a detailed account of the distribution of endemic or rare species that comprise part of the chasmophytic and rupestral vegetative community lining the Maltese coastal cliffs. The species discussed in this paper are the following: Palaeocyanus crassifolius (Bertoloni) Dostál; Cremnophyton lanfrancoi Brullo and Pavone; Helichrysum melitense (Pignatti) Brullo, Lanfranco, Pavone and Ronsisvalle; Linaria pseudolaxiflora Lojacono; Asparagus horridus L. and Allium sphaerocephalon subsp. arvense (Guss.) Arcang. The first four are species found in article 17 of the Maltese habitats and species, listed in the annexes of the Habitats Directive by the European Commission. Asparagus horridus and Allium sphaerocephalon subsp. arvense are recent additions to the flora of Malta, and their distribution is given in this account. Discussion on the distribution and observations on the preferred habitat are given for each species supplemented by recommendations for the red data book.
Keywords Article 17, coastal cliffs, endemic, Malta, Maltese flora, rupestral species
Citation Mifsud, S. (Aug 2013): Distribution of some rare or endemic chasmophytic and rupestral species growing along the coastal cliffs of the Maltese Islands .   Webbia: Journal of Plant Taxonomy and Geography, Vol.68:1: pp35-50, US. (DOI: 10.1080/00837792.2013.807451)
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Paper 18 Althenia orientalis (Tzvelev) Garcia-Mur. & Talavera (Zanichelliaceae Dum.): An addition to the Maltese Flora (Central Mediterranean)
Author(s) / Yr. of Publ. Alex Casha & Stephen Mifsud (Aug 2013)
Abstract Althenia orientalis (Tzvelev) García-Mur. & Talavera has been recorded from brackish to saline waters at the Ghadira Nature Reserve in Mellieha, Malta, and constitutes a new species for the Maltese flora. Details on the taxonomy of the species and the habitat of the single population are conveyed, while the local morphology and introduction in Malta of this hydrophyte are further discussed
Keywords Keywords, Zannichelliaceae, Althenia, halophyte, hydrophyte, Mediterranean flora, Malta
Citation Casha, A. & Mifsud, S. (Aug 2013): Althenia orientalis (Tzvelev) Garcia-Mur. & Talavera (Zanichelliaceae Dum.): An addition to the Maltese Flora (Central Mediterranean) .   Webbia: Journal of Plant Taxonomy and Geography, Vol.68:1 pp57-61, US. (DOI: 10.1080/00837792.2013.807452)
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Paper 19 Preliminary observations from long-term studies of Gladiolus l. (Iridaceae) for the Maltese Islands
Author(s) / Yr. of Publ. Stephen Mifsud and Anthony P. Hamilton (Aug 2013)
Abstract Field studies in 28 random sites in Malta and Gozo were carried out between 2010 and 2012 to investigate Gladiolus species on the Maltese islands. The results obtained were significantly very different from the general picture of Gladiolus spp. reported for several decades for the Maltese islands, which inferred that the populations were mainly of G. italicus, with unsubstantiated records of G. byzantinus (a garden cultivar) and G. dubius (a doubtful and now unrecognized species). The results from these studies clearly show the abundant presence of G. communis, but the rather rare G. italicus, the latter not being observed in more than half the sites that have been surveyed. This paper deals with an historical account of Gladiolus spp. in Malta, an historical account and the origins of G. italicus, a taxonomic distinction between the two species confirmed from Malta and a discussion based on their seeds and origins.
Keywords Iridaceae,Gladiolus, flora, Maltese Islands
Citation Mifsud, S. & Hamilton, P. (Aug 2013): Preliminary observations from long-term studies of Gladiolus l. (Iridaceae) for the Maltese Islands.   Webbia: Journal of Plant Taxonomy and Geography, Vol.68:1 pp51-56, US. (DOI: 10.1080/00837792.2013.779817)
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Paper 20 Polypodium vulgare L. subsp. melitense new subspecies from Gozo, Maltese Islands (Pteridophyta: Polypodiaceae Bercht. & J. Presl.)
Author(s) / Yr. of Publ. Adalberto Peroni, Gabriele Peroni & Stephen Mifsud (2013)
Abstract Polypodium vulgare L. subsp. melitense subsp. nov. from the island of Gozo (Malta) is described. Polypodium vulgare s.l. is being recorded for the first time from Malta. Comparison of the new subspecies with other Polypodium taxa present in continental Europe and the Mediterranean area is outlined. Pteridophyta of the Maltese Islands are finally listed
Keywords Pteridophyta, Polypodiaceae, Polypodium vulgare new subspecies, Gozo, Malta.
Citation Peroni, A., Peroni, G. and Mifsud, S. (2013): Polypodium vulgare L. subsp. melitense new subspecies from Gozo, Maltese Islands (Pteridophyta: Polypodiaceae Bercht. & J. Presl.) .   Botanica Complutensis, 37: 41-46. 2013. ISSN: 0214-4565
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Paper 21 Recent records of spiders (Arachnida: Araneae) from Malta
Author(s) / Yr. of Publ. David Dandria, Alfred Gatt Floridia & Stephen Mifsud (2014)
Abstract Two spider species are recorded in Malta, one for the first time (Zora sp. ?spinimana, Fam. Zoridae) and one for the second time (Aphantaulax cincta, Fam. Gnaphosidae). Observational data and other information about the species together with photographs are provided.
Keywords "Zora spinimana", "Aphantaulax cincta", Malta, Araneae
Citation Dandria, D., Gatt Floridia, A. & Mifsud, S. (2014): Recent records of spiders (Arachnida: Araneae) from Malta.   The Central Mediterranean Naturalist, Vol.5, Pts. 3 & 4 (2011-2012), pp:6-8, Nature Trust, Malta
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Paper 22 Marasmius corbariensis (Roum.) Singer (Family Marasmiaceae Roze Ex Kuhner 1980) – A new fungal species for Malta
Author(s) / Yr. of Publ. Stephen Mifsud (2014)
Abstract A population of Marasmius corbariensis (Roum.) Singer was found in an olive grove at Xewkija, Gozo. Being a new agaric for the Maltese islands, details on this population and habitat is given in this communication.
Keywords Marasmius corbariensis, Agaricales, Marasmiaceae, fungi, Malta, new record
Citation Mifsud, S. (2014): Marasmius corbariensis (Roum.) Singer (Family Marasmiaceae Roze Ex Kuhner 1980) – A new fungal species for Malta.   The Central Mediterranean Naturalist, Vol.5, Pts. 3 & 4 (2011-2012), pp:52-53, Nature Trust, Malta
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Paper 23 Bromus cathatrticus Vahl (Fam. Poaceae), A new record for the Maltese Islands
Author(s) / Yr. of Publ. Timothy J. Tabone, Alex Casha & Stephen Mifsud (2014)
Abstract First records of Bromus catharticus Vahl from Malta are given in this short communication. A short account of the the naturalising potential and sometimes the invasiveness of this species is given. Notes on the local populations are given and routes of possible introduction are discussed.
Keywords Alien flora, Bromus, Malta, Poaceae, New record
Citation Tabone, T.J., Casha, A. & Mifsud, S. (2014): Bromus cathatrticus Vahl (Fam. Poaceae), A new record for the Maltese Islands.   The Central Mediterranean Naturalist, Vol.5, Pts. 3 & 4 (2011-2012), pp:65-67, Nature Trust, Malta
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Paper 24 A study of the genus Persicaria Miller (Polygonaceae) in the Maltese Islands
Author(s) / Yr. of Publ. Stephen Mifsud (2014)
Abstract A detailed study on the taxonomy, distribution and populations of Persicaria spp. occurring on the Maltese Islands is carried out based on field surveys between 2008 and 2011. Four taxa are recognised in this study: two forms of Persicaria senegalensis, P. lanigera and P. salicifolia, of which only the latter is native. A taxonomical overview of these species and a detailed account of the distribution and size of the populations of each species (including new records) is given. Habitat preference of the species and their significance in Maltese wetland ecosystems are discussed.
Keywords Persicaria, Polygonum, Taxonomy, Distribution, Wetland flora, Maltese Islands
Citation Mifsud, S. (2014): A study of the genus Persicaria Miller (Polygonaceae) in the Maltese Islands.   The Central Mediterranean Naturalist, Vol.5, Pts. 3 & 4 (2011-2012), pp:26-51, Nature Trust, Malta.
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Paper 25 Chromosome number of Elatine gussonei (Sommier) Brullo (Elatinaceae) and its distribution on the Maltese Islands
Author(s) / Yr. of Publ. Anna Kalinka, Stephen Mifsud, Agnieska Popiela, Magdalena Achremi (2014)
Abstract Elatine gussonei (Sommier) Brullo is an endemic species, with a distribution restricted to the central part of the Mediterranean Basin (Maltese islands, Lampedusa, southern part of Sicily). This hydrophyte grows in rainwater pools and cavities in karstic limestone. Although the morphology has been well studied, no karyological study has been carried out, and hence this work brings the first chromosome data for the Maltese-pelago endemic E. gussonei.We have found a diploid number of 54 chromosomes in E. gussonei, which differs from the chromosome number of most of Elatine species (2n = 36). Additionally, this account gives a recent distribution of the species on the Maltese islands.
Keywords Elatine gussonei, chromosome number, phytogeography
Citation Kalinka, S. et al. (2014): Chromosome number of Elatine gussonei (Sommier) Brullo (Elatinaceae) and its distribution on the Maltese Islands.   Acta Botanica Croatia, 73 (1), 267-273, Croatia.
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Paper 26 Promotion of the Maltese Ophrys bombyliflora forma parviflora to a variety
Author(s) / Yr. of Publ. Stephen Mifsud (Dec 2014)
Abstract A discovery of a second clump of Ophrys bombyliflora f. parviflora from the Maltese islands was found to be formed of a homogenous population, the same previously found in 2008. These micro-satellite populations show a degree of genetic barrier with the normal form of O. bombyliflora, despite they are in sympatric associations. In addition, on evaluating its distinguishing characters, and abiding with the current application of infraspecific ranking in plant taxonomy, this taxon is here combined to a variety level, hence Ophrys bombyliflora var. parviflora.
Keywords Orchids of Malta, Flora Malta, Ophrys bombyliflora var. parviflora, endemics.
Citation Mifsud, S. (Dec 2014): Promotion of the Maltese Ophrys bombyliflora forma parviflora to a variety.   Journal Europaischer Orchideen , 46 (3/4): 2014. 679-685, Germany.
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Paper 27 Two hybrid orchids described from Malta: Ophrys x tumentia nothosubsp. tumentia and Ophrys x tumentia nothosubsp. sancti-martinii
Author(s) / Yr. of Publ. Stephen Mifsud (Dec 2014)
Abstract Mifsud, S. (2014): Two hybrid orchids described from Malta: Ophrys x tumentia nothosubsp. tumentia and Ophrys x tumentia nothosubsp. sanctimartinii.- J. Eur. Orch. 46 (3/4): 686-700. Two hybrid orchids of Ophrys caesiella with, respectively, the very early flowering Ophrys iricolor subsp. hospitalis and the later flowering Ophrys iricolor subsp. vallesiana are described from Malta. The descriptions of these two hybrids are supported by respective intermediate characters found flowering in sympatric association with their parent orchids.
Keywords Orchidaceae; Ophrys caesiella; Ophrys iricolor subsp. hospitalis; Ophrys iricolor subsp. vallesiana; Ophrys x tumentia nothosubsp. tumentia; Ophrys x tumentia nothosubsp. sancti-martinii; Malta.
Citation Mifsud, S. (Dec 2014): Two hybrid orchids described from Malta: Ophrys x tumentia nothosubsp. tumentia and Ophrys x tumentia nothosubsp. sancti-martinii.   Journal Europaischer Orchideen, , 46 (3/4): 2014. 686-700, Germany.
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Online Articles of Scientific Value


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Article 01 A FLORISTIC SURVEY ON THE GOZITAN ISLETS OF TAC-CAWL AND TAL-HALFA IN THE MALTESE ISLANDS
Author
(Yr. of Publ.)
Stephen Mifsud (2011)
Abstract A floristic survey was carried out on the islets of tal-Halfa and tac-Cawl, situated at the Southern coast of Gozo, to confirm or not the results of a recently published paper where it was concluded that a number of species from these sites are extinct or doubtful from a preceding work carried out by a group of scientists in year 2000. This paper confirms the occurrence of most of the species found by the latter scientists and additionally reports a significant number of new records of vascular plants of which some are legally protected, listed in the national red data book or considered rare. Taxonomical notes about few ambiguous plants that are found are also discussed.
Keywords tac-Cawl / tal-Halfa / Islets / Malta / Gozo / Flora
History Version: 1.0 released on 03-Jun-11 (First release)
Version: 1.1 released on 05-Jun-11 (Minor changes)
Version: 2.0 released on 15-Nov-11 (Revisit at tac-awl rock in Autumn (Nov 2011))
Version: 3.0 released on 5-Dec-11 (Revisit at tal-Halfa rock in Autumn (Dec 2011))
Version: 3.1 released on 30-Aug-13 (Proof-read by Brian Farrugia)
Citation Mifsud, S. (2011): A FLORISTIC SURVEY ON THE GOZITAN ISLETS OF TAC-CAWL AND TAL-HALFA IN THE MALTESE ISLANDS.MaltaWildPlants.com Online Publications; Ref: MWPOP-001. Article Update Version: 3.1 released on 30-Aug-13
URL Link: http://www.MaltaWildPlants.com/publ/index.php#W01
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